Palace's development side saw their three game winning streak come to an end at the hands of Crewe Alexandra at Tooting and Mitcham's KNK Stadium.
After falling behind early on Gary Issott's side drew level thanks to Ryan Inniss' header, but the visitors took the lead once again on the hour mark and this time there wasn't to be another Eagles equaliser despite their best efforts.
Missing the likes of Jake Gray, Hiram Boateng and Sullay Kaikai, a younger-than-usual home side took to the field but only four minutes had elapsed when the visitors found a way to edge in front. Callum Saunders got past Rodigue Ndiane on the left flank to reach the byline, and his cutback was rifled home first-time by James Jones, beating Chris Kettings at his near post.
That seemed to deflate Issott's side ho found chances few and far between during the opening exchanges with only a Morgan Ferrier shot that was hit straight at Scott Shearer all they could muster, but the Railwaymen continued to threaten and Greg Leigh should have done better with a header from Ollie Turton's cross but missed the target.
Just after the half-hour mark Palace had a great chance to take the lead when Jacob Berkley-Agyepong whipped a dangerous free-kick into the box which Inniss steered wide, but two minutes later he made no mistake when he rose highest to nod Reise Allassani's corner into the ground and past Shearer to equalise.
That lifted the Eagles and five minutes before half-time Ferrier embarked on a powerful run into the final third before unleashing a shot from 25 yards that beat Shearer but cracked against the crossbar.
After the restart Michael Chambers met a Berkley-Agyepong corner and what looked like a routine save for Shearer nearly slipped through his grasp, but he recovered to start a quick counter-attack that saw Kettings called into action at the other end, blocking Brad Imman's effort with his feet.
From the resulting corner Kettings' punch fell to Chris Atkinson who hammered the ball goalbounds with the keeper stranded on the ground, but he was saved by Mandela Egbo's goalline clearance.
But on 65 minutes Crewe did find themselves back in front in dubious circumstances. A long searching ball came into the box which found an offside-looking George Cooper, but with the Eagles defence waiting for the linesman to flag it never arrived and the striker slammed the ball home from from close range.
Searching for a second equaliser, Egbo saw a low curling effort just miss the bottom corner of Shearer's goal and his fellow full-back Rodigue Ndiane forced the Crewe keeper to push a stinging drive over his bar but ultimately their unbeaten run came to an end.
Palace: Kettings, Ndiane, Egbo, Sow (Black 19), Inniss, Chambers, Allassani, Dymond, Ferrier, Adarabioyo (Andrews 72), Berkley-Agyepong (Coker 72). Subs not used: Croll, Magri
Crewe: Shearer, Baillie, Leigh, Mullarkey, Na, Turton (Johnston 74), Jones, Atkinson, Saunders (May 70), Cooper, Inman (Kearns 88). Sub not used: Howell.